And yet they're 'working for' the Garleans, who are known to kill beastmen whether they will summon a Primal or not. Either there's a plot-hole, Varis zos Galvus (who wanted to kill the Zundu Vanu Vanu) is lenient to his sons decisions, or Lupin aren't the typical beastmen we're familiar with (compared to others, they're the most 'human').