They are both shown in different lights, this makes a HUGE difference.
By skin-tone he probably meant skin definition, how clear it is that its actually skin (Not just a solid colour)
The difference isn't that phenomenal from the two screenshots he's shown really.
Personally, I don't see the fuss. This game looks great so far considering how much they're having to limit it.

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yea, the top one did look a little cel-shaded.



