I think we probably need to hear from Yoshi-P, since it's clear a lot of the problems are definitely goof ups, so maybe most of the difference are. Like the moonkeeper fangs are probably a mistake and not on purpose, but if they can screw something like that up maybe they screwed up some other things like some lips.

It would be really bizarre if they said all that stuff about not wanting to ruin anyone's image of their character, only to then completely donk up like 1 in every 6 faces on purpose.