Hildibrand
The punchline to the "Mandervilles are descended from aliens" bit is that Delion, alien conspiracy theorist extraordinaire, is completely unfazed by this. You'd think he'd go into a rant, but instead he just says something to the effect of "Figures." and moves on.
But yes, between Godbert's idle musings and Nashu pointing out the bit doesn't explain Julyan, it's implied the "explanation" is being set up as a punchline in the last quest. Something something Godbert was adopted, or whatever.
I don't mind MSQ Lore supporting Hildibrand, because he's so over the top it's impossible to backport much of the latter into the former. About the only thing from Hildibrand they've actually used in the MSQ is Godbert (albeit acting in a much more subdued manner) and Hancock referencing the Heavensward questline ("Stealer of Pants").
Nashu playing basketball with a coblyn, bricking, and then the PC dunking said coblyn into Lugae was definitely the highlight. Coming up with the hasty excuse of "I have an Orchestrion addiction. Spent my last gil on Return to Oblivion," the city guards thanking Hildibrand for using his stupidity and poor judge of character to sus out a shady merchant, and Brandihild's obsession with the low-poly Labyrinthos grapes were all very funny as well.