Ability button dot crawl meaning?

[Darkstride]

Thanks. I searched all over, and the only thing I could find about combo actions was articles on how to macro them, and nothing about what the tooltips actually meant.
And,another item if anyone can answer. I ran a dungeon yesterday, and Aeolian Edge was greyed out the whole time. Couldn't use it at all.
Now that I think of it, the dungeon was level 24, so perhaps I was level synced to it, and lost any abilities that came later?

Yeah, you level synced and lost access to it.