Per the official blog: "These can only be used in Pyros and the next iteration of Eureka, but you'll be able to use them as soon as you enter Pyros!"
https://na.finalfantasyxiv.com/pr/blog/



Per the official blog: "These can only be used in Pyros and the next iteration of Eureka, but you'll be able to use them as soon as you enter Pyros!"Other things still on the table:
I've read speculation that Pyros might not be the final part of Eureka... we'll know more after 4.55 drops. In the previous expansion HW, the relics final step was equal in ilvl to the 8-man raid weapons. So if the Relic doesn't get to i405 in Pyros, it's possible that a final area will be implemented for the final step...
It's doubtful to me that we'll get a "victory lap" of content outside of Eureka for the 4.0 Relic, but I guess they could do a Victory Lap that requires you to go through all the Eureka areas and kill specific FATE targets with your 4.0 Relic equipped... /pray for those that have to defeat Cassie... If such a Victory Lap is a step in the 4.0 relic, then a 4th part of Eureka would be unlikely.
..
https://na.finalfantasyxiv.com/pr/blog/
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