Interesting subject but I just wanted to add some more facts about Ramuh that the dailies have given us.
Ramuh at least to the Slyphs isn't so much summoned as called from the heaves. While it is vague it is possible that Ramuh is actually out of the aether and roaming the sky above Eorzea. But that would beg the question why would he leave tempered followers about and then leave them to their own devices? Well Slyphs unlike the other beastmen appearently require a certain Slyph to call upon him. Not every Slyph can do it, so its not so much the need for lightning crystals but the need for the special slyph to be given reason to Call him, and by how the quest is read Control him. This special slyph like all slyph are grown and then become slyph from special conditions and in the quest you hijack the special one before the birth to avoid the tempered ones to temper the special one. So what that points to if this isn't reconned is that the tempering was a request made by the last special slyph to control Ramuh and a method to temper to Ramuh with out Ramuh present was created. So effectively it was one bad seed (pun) that was uncharacteristically violent or fearful that lead to the inital temperings not specifically Ramuh.
Also from the main story line the kobold seem to summon titan in response instead of proactively. But you could make that argument about the Ixail as they original took up specific grudges after loosing access to Twelveswood and the lost of their feathers/the ability to fly. The only beastmen that seem warlike from introduction are the Amaljaa and the sahagian both of which summoned or attempted to summon their primal as a armament for their new enemy. In this case 1.0 Uldah because they choice to abandon them and "remove" them under threat of Garlemand and 2.0 Limsa part of the island that was sahagian is no longer claimed by the sea via the dalamud fall. Both of which you could claim are reactive as they did not choose the battle placed before them. But in theory neither did the dragon that made its home in Brayflox or in Stone Vigil as we haven't seen any notion that the dragons have any intelligence beyond that of a wolf beside Nidhogg who seems not only intelligent but sentient.

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