I read through (most) of the first five pages, the other five got a little tl:dr...
This is all entirely IMO...
I remember (once upon a time) googling how cats mated. The tom-cat'spenisis barbed, which causes pain for the girl when he pulls out. This stimulates her to ovulate, which explains why cats never have periods, like dogs and humans do (totally not suggesting they can cross-breed).
What does this mean? Well, maybe Miqo'te men and women are similar? Which can also explain why there are so many Miqo'te prostitutes, and are able to get away with it (as in, not having kids), because only a male Miqo'te can 'stimulate' the female to ovulate(unless someone has invented a way...).
For Lalafells... They're just too small. A male Roe and a fem Lala? Honestly, how will that work? If it were a male Lala, there'd really be only one getting anorgyoutta that, unless he had is entire lower half in her. If that produces a kid... I'd be scared.
All in all, I can only see Hyurs, Elezen and Roegadyn being able to cross-breed. The Miqo'te and Lalafells keeping to their own.
As far as the half'n'half kids being rare, I agree that it is a cultural thing. Look at America. There's pretty much every culture in the world there, and how many do you see intermingling and making babies? It's probably a lot more common in America, but it's still not that common.
Hyurs and Elezen founded Gridania. I'm sure a handful did make human/elf babies, but it's not common enough that Hyurs and Elezen "purebreds" became minority.
Infertility? A good possibility. Male mules (horse + donkey) are sterile and something like 5% of the female mules can reproduce.

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